The round was led by Remiges Ventures with participation from OrbiMed, Dementia Discovery Fund, Arix Bioscience, RMGP Bio-Pharma investment Fund and others.
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Fast normal- feathering one-day-old chicks have six primary remiges and at least another six secondary remiges (see Figure 3A). While "retarded" tsts one-day-old chicks have normal primary remiges but only the first three (external) secondary remiges. At 10 days of age, differences between normal and "retarded" chicks are even more evident. "Retarded" chicks show the first three or four primary remiges of normal length, with each new feather being shorter than the preceding one (see Figure 3B).
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At three weeks of age, "retarded" chicks have even fewer primary remiges than normal chicks, and only the first three secondary remiges reach the same length as primary remiges. Figure 3. Effect of autosomal genes of T series on 10-day-old chicks. (A) Normal-feathering, with a T− genotype.
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"Tardy" tt chicks are more abnormal than "retarded" chicks. They show normal primary remiges at hatch, but lack secondary remiges. At ten days of age they still lack rectrices and show small or no development of secondary remiges (see Figure 3C). Rectrices do not appear until at least eight weeks of age.
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Feathering types Diagram of the wing of a one-day-old chick in dorsal view: A fast normal-feathering with the primary remiges (1a) longer than the coverlets (2a). B sex-linked delayed-feathering with the primary remiges (1a) of the same length than the coverlets (2a). C modified delayed-feathering with the primary remiges. (1a) shorter than the coverlets (2a).
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It is similar to the dusky grasswren, though the markings are generally brighter and more defined, e.g. the wing coverts and remiges are grayish with a small rufous patch at the base of the outer remiges, compared with the same parts of the dusky grasswren being a uniformly dull rufous-brown.
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This is a short-tailed robust bird with a long hook-tipped bill; like other tityras it has a peculiar vestigial ninth primary feather. The adult black-tailed tityra is about long and weighs about . The male is dull white above and white below. The rectrices, the primary and secondary remiges and a cap extending to below the eyes are black; the tertiary remiges are silvery grey.
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Lores, rump and underside rusty. Wing coverts with some rusty edging. Remiges with paler inner surfaces. Underside of wing dusky brown with paler edges to coverts.
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Remiges are dark brown with yellowish edges to secondaries, forming a yellow-olive panel when the wing is folded. Uppertail is olive-brown with yellow-olive outer edges. Underbody is mainly light brown- grey, with pale yellow streaks in the centre of the breast, pale yellow on the upper belly, flanks and undertail coverts, and cream on the lower belly. Underwing coverts are off-white with brown-grey remiges.
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The rectrices and the secondary and tertiary remiges are pure grey, the latter contrasting with the brown forewing and the black primary remiges at the wingtips. The upperside and underside of the wing look similar, though the brown is lighter on the underwing. Whether from the side or below, flying males appear characteristically three-colored brown-grey-black. The legs, feet, irides and the cere of the black bill are yellow.
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It has sometimes been considered a subspecies of the pied imperial pigeon, but has a yellowish tip to the bill, black spotting near the vent, and silvery-grey remiges.
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There are some multifactorial autosomal genes which affect only chicks with delayed-feathering and have no effect on fast normal-feathering chicks, nor on adult plumage. The sole effect of multifactorial slow-feathering is to reduce the length of the primary remiges of one-day-old chicks without a concomitant reduction of the primary coverlets.Poultry Science. 55, 2094 (1976) That is, one-day-old chicks with primary remiges shorter than coverlets carry the K gene of sex-linked delayed- feathering (see Figure 2C).
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Migrating bird in Cordell Bank National Marine Sanctuary offshore California, United States; note upperwing pattern Adults birds are 46–47 cm in length, with a 97– to 99-cm wingspan, and have been recorded to weigh 342-425 g. The upperside of Buller's shearwater is bluish grey. A blackish stripe runs from the tertiary remiges to the primary wing coverts. The primary remiges are blackish, also; the two black areas do not meet at the hand, however; the area between them is a rather light grey, and under bright light may appear almost white.
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This strong attachment is accomplished by ligaments under the skin, which in some birds and other feathered dinosaurs results in raised bumps or marks along the rear forelimb bone (ulna). These bumps, called quill knobs (ulnar papillae), are often used as an indirect indication of strongly-attached forelimb feathers in fossil species, and can also indirectly indicate the number of secondary remiges in a given specimen. Flight feathers (remiges and rectrices) are specialized types of pennaceous feathers, adapted for high loadings and often strongly asymmetric for improved flight performance.
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Adults are strikingly tri-colored, with a black back and tail and a red head and neck. Their underparts are mainly white. The wings are black with white secondary remiges. Adult males and females are identical in plumage.
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Wing and tail flight feathers - rectrices and remiges - are between red and red-violet. The wing covert feathers are black and red with black tips. The leg feathers are bluish-purple. The feet are medium gray, and the claws are black.
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The scapulars (shoulder feathers) are white, and the wings are black with a white bar made up by the bases of the primary remiges, continuing slightly offset onto the bases of the secondary remiges in some regions. The tail is black, long, and pointed at the tip; the outer rectrices have white outer vanes. The underparts are white, slightly tinged with grey in most subspecies. In particular the breast is usually darker and sometimes browner than the rest of the light underside, and may appear as an indistinct band between the lighter belly and white throat.
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Life restoration by Emily Willoughby The skeletal morphology of Serikornis suggests a terrestrial ecology without flying adaptations. The tail is covered proximally by filaments and distally by fine rectrices. Symmetrical, barbule- free remiges are attached along the forelimbs and elongated feathers of the hind limbs extend to the toes, suggesting that the remiges of the hind legs had evolved in the maniraptorans residing on the ground before being co-opted to an arboreal lifestyle and possibly, gliding. The coracoid of Serikornis is devoid of diagnostic ornamentation present in Anchiornis but is distinguished by having the tuberculous coracoid elongated to form a crest.
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The second metacarpal of the metacarpus of the hand, the bone that primary remiges attach to, also had a flat bony shelf as its dorsal surface. The shelf made a perfect spot for the primary feathers to lay across in their life-attachment.
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The juvenile has less contrast in the wings, but the remiges bear prominent white spots. It differs from greater spotted eagle juveniles by a lack of wing covert spotting and the presence of a cream-colored neck patch. The call is a dog-like yip.
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Canada goose flying Modern pennaceous feathers and remiges are known in the advanced maniraptoran group Aviremigia. More primitive maniraptorans, such as therizinosaurs (specifically Beipiaosaurus), preserve a combination of simple downy filaments and unique elongated quills.Xu X., Zheng X.-t. and You, H.-l. (2009).
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Due to a high-calorie diet, especially one high in proteins and/or low in vitamin D, vitamin E, and manganese, one or both carpus (wrist) joints are delayed in their development relative to the rest of the wing; for reasons unknown, if only one wing is affected, it is usually the left one. The result is a wrist which is twisted outwards and unable to perform its usual function. Angel wing symptoms include stripped remiges (flight feathers) in the wrist area, or remiges protruding from wings at odd angles. In extreme cases, the stripped feathers may resemble sickly blue straws protruding from wings.
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In immature birds, both male and female chicks have remiges, scapulars and wing coverts intermixed with buff, outer rectrices infused with white and feathers more narrow and pointed than adults. Immature females have a brown breast. Trogons are the only birds with a heterodactyl toe arrangement.
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The sharply two-toned basic pattern of the rufous-tailed flatbill - an olive-green bird with a bright rufous tail and wings - is distinctive and easily identifiable. Compare to dusky-tailed flatbill (Ramphotrigon fuscicauda), which has dusky rectrices and remiges and a more heavily streaked breast.
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It forms the tip of the wing skeleton in birds. To it, most of the primary remiges attach. The alula, by contrast, is formed by the thumb, which does not completely fuse with the other hand-bones. Likewise, the tipmost primaries attach to the phalanx bones.
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The upper body is a dark greyish- brown to olive-brown. Olive-green outer edges on the remiges combine to form an olive panel on the folded wing. The bill is black and slightly down-curved, and the gape is cream. The legs and feet are grey-brown.
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Apart from the difference in size, males and females look exactly alike. Immature birds have light grey plumage with darker brownish nape and remiges. Their bare parts are dull grey. Chicks are naked at first, then grow white down feathers all over, before moulting to the immature plumage.
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Eyes rounded, yellow to orange with black pupil and lightly framed by a thin, flesh-colored eye ring. Back, rump and wings brownish gray, which in adult males with metallic blue shoulders. The primary and secondary remiges are blackish brown. The chest, blue-dark gray, stained purple in males.
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Cassowaries have small wings with 5–6 large remiges. These are reduced to stiff, keratinous quills, resembling porcupine quills, with no barbs. A claw exists on each second digit of the feet. The furcula and coracoid are degenerate, and their palatal bones and sphenoid bones touch each other.
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K is another dominant allele of the allelic series which controls feather growth rate. One-day-old chicks have primary remiges and coverlets of the same length (see Figure 2B). In 8- to 12-day-old chicks rectrices are not yet developed (see Figure 1, chick on the right). Figure 2.
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Serrated edges along the owl's remiges bring the flapping of the wing down to a nearly silent mechanism. The serrations are more likely reducing aerodynamic disturbances, rather than simply reducing noise. The surface of the flight feathers is covered with a velvety structure that absorbs the sound of the wing moving.
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Males in nonbreeding plumage have largely white underparts, supercilium, and cheeks. The upperside feathers have brown fringes, and most wing feathers white ones, giving a scaly appearance. The bases of the primary remiges are also white. The coloration renders the adult male rose-breasted grosbeak (even while wintering) unmistakable if seen well.
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A small green iridescent patch is present on the outer scapular feathers, besides, especially in the western race, the sides of the lower breast. Wing coverts and remiges are bordered in a paler or warmer tone. The sexes are similar. Immature birds are dun brown above with buff head and underpart plumage.
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It also has a dusky lore. The throat is whitish and the chest is a pale gray with inconspicuous dusky streaking, while the belly and undertail coverts are a pale yellow. Additionally, this species has dark brown, well-rounded wings with pale cinnamon-edged coverts and remiges. The wings are about long.
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The other measurements are either from the freshly killed bird or are unlikely to have changed. Dull dusky black overall, with lighter brown feather edges which are prominent on the body feathers and less conspicuous on the remiges and tail. Iris yellow. Feet dusky brownish; bill the same colour or somewhat lighter.
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Adults are long. They have long pink legs, a long thin black bill and are blackish above and white below, with a white head and neck with a varying amount of black. Males have a black back, often with greenish gloss. Females' backs have a brown hue, contrasting with the black remiges.
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When it finishes its moult migration, this bird moults its remiges between August and September, which makes it unable to fly. This moult is preceded by an increase in weight. During the moult, the breast muscles atrophy. When the moult is completed, this grebe continues to gain weight, often more than doubling its original weight.
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Handbook of the Birds of the World. Vol. 8. Broadbills of Tapaculos. Lynx Edicions, Barcelona. The male has black wings with grey edging to the remiges (sometimes very narrow, and barely visible in the subspecies where the males are primarily black), and white wing-bars that may appear spotty, especially on the lesser wing coverts.
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The crown is glossy black in males. The nape is pale grey with a grey- white to white band separating the crown from the back. The wings are dusky greyish brown with white edging on the inner remiges and wing-coverts. The tail is dusky brown, and the stout bill, iris and legs are black.
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The male of the species has a black hood, mandible, and throat, as well as a black tail. Wings are black, but the remiges and rectrices (flight feathers) are fringed with white. The secondary coverts form yellow epaulets. The back and vent are yellow washed with olive, and the underside is almost uniformly yellow.
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These feathers are not completely renewed until the beginning of October. The molt of the tertiary remiges begins more or less at the same time as that of the secondary ones. The middle tertiary falls first, followed by the internal tertiary. Both feathers are often well developed again before the outer tertiary leaves the sheath.
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The mantle, back and wings are olive-greenish, and usually spotted or barred in buffy to golden yellow. The shafts of the remiges and rectrices are yellow to golden yellow. The underpart plumage is spotted black to a lesser or greater degree. Some species include drumming on dead wood as a means of non-vocal signaling.
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All have white under- and upper-tail coverts, and several have some extent of white on their heads. The neck, body and wings are grey or white, with black or blackish primary—and also often secondary—remiges (pinions). The closely related "black" geese in the genus Branta differ in having black legs, and generally darker body plumage.
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The fulvescens form of the greater spotted eagle must be distinguished from the pale forms of the tawny eagle via their underwing pattern, often with completely blackish underwing coverts and usually plain looking dark remiges over the entire primaries with more distinct pale carpal arcs.Davidson, I. (1978). Flight identification of southern African raptors. Bokmakierie, 30: 43-48.
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CRC Handbook of Avian Body Masses by John B. Dunning Jr. (Editor). CRC Press (1992), . They typically have strong, stubby beaks, which in some species can be quite large; however, Hawaiian honeycreepers are famous for the wide range of bill shapes and sizes brought about by adaptive radiation. All true finches have 9 primary remiges and 12 rectrices.
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It was estimated that a complete series might include fifteen of these papillae ulnares. The ulna is long and the other lower arm bone, the radius, measures . The hand bones show that their joints allowed for little mobility. The wingspan of Dakotaraptor was estimated at , not taking into account possible primary remiges longer than the hand.
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Elderly macaws show a ring of dark grey surrounding the pupil where the iris has become thinner and the back of the retina shows through. It can be separated from the slightly larger blue-and-yellow macaw by the blue (not black) throat, the blue (not green) crown and the lack of contrast between the remiges and upperwing coverts.
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Total length c. . As other members of the Pyrrhura picta complex, it is a long-tailed mainly green parakeet with a dark red belly, rump and tail-tip (tail all dark red from below), a whitish or dull buff patch on the auriculars and bluish remiges. The cheeks and ocular region are dark maroon. The nominate subspecies (P. a.
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Later he was called also tribunus liburnarum (tribune of warships). The crew consisted of the officers (trierarchus), the rowers (remiges) and a centuria Marines (manipulares/milites liburnarii). The team (classiari/classici) were divided into two groups, the technical staff and the Marines. The service was 26 years (as opposed to the 20 to 25 years for a legionary).
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Bentsen State Park, Texas, US Both males and females have a black mandible and throat, as well as a black back and long black tail. Wings are black, but the remiges and rectrices (flight feathers) are fringed with white. These form a single white wing bar and white wing spots when folded. The secondary coverts form orange epaulets.
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The mantle, back and wings are olive- greenish, and usually spotted or barred in buffy to golden yellow. The shafts of the remiges and rectrices are yellow to golden yellow. The underpart plumage is spotted black to a lesser or greater degree. Some species include drumming on dead wood as a means of non-vocal signaling.
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The underwing coverts are dark brown, with the remiges being pale streaked with black, overall imparting the wings of adults a dark look. The underside of the tail has similar barring as the remiges while the upperside is the same uniform brown as the back and upperwing coverts. The eyes of mature martial eagles are rich yellow, while the cere and large feet pale greenish and the talons black. Martial eagles have a short erectile crest, which is typically neither prominent nor flared (unlike that of the crowned eagle) and generally appears as an angular back to a seemingly flat head. This species often perches in a quite upright position, with its long wings completely covering the tail, causing it be described as “standing” rather than “sitting” on a branch when perched.
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The male of the subspecies aspersiventer of the Andes in north- western Bolivia and adjacent Peru approaches melanochrous in colour, but has dense white barring to the belly. male Thamnophilus c. melanochrous. Females are equally variable. In most of this species range, their wings are blackish- brown with rufescent edging to the remiges, and white or buff wing-bars (often appears rather spotty).
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South Island takahe (Porphyrio hochstetteri) from behind, showing the short, soft, and fluffy remiges typical of flightless rails The rails are a family of small to medium-sized, ground-living birds. They vary in length from and in weight from . Some species have long necks and in many cases are laterally compressed. The bill is the most variable feature within the family.
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The wingspan of adult geese is . At Moscow Zoo The upperparts are greyish-brown, with thin light fringes to the larger feathers and a maroon hindneck and cap (reaching just below the eye). The remiges are blackish, as are the entire underwing and the white-tipped rectrices, while the upper- and undertail coverts are white. A thin white stripe surrounds the bill base.
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The Kimberley honeyeater is similar in appearance to the white-lined honeyeater, having dark grey upperparts, light grey underparts, grey eyes, with dark grey below the eyes and a black beak. It is distinguished from the white-lined honeyeater by the lack of citrine edging on the upper surface of the remiges and , pale creamy-buff , and a milky-white belly.
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It is somewhat bent to the inside. The most notable anatomical feature is the row of very prominent bumps along a ridge on the lower edge of the ulna, one of the forearm bones. These are called ulnar papillae, or quill knobs. In birds and some other theropod dinosaurs, these bumps were spots for reinforced attachment of the remiges, or wing feathers.
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Ks ("slow") is another dominant allele of the allelic series which controls feather growth rate. In one-day-old chicks primary remiges are shorter than the coverlets. Plumage growing is greatly delayed during the first weeks of juvenile life, but birds of both sexes completely finish plumage growing at twelve weeks of age. This allele has no effect on adult plumage.
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The undertail coverts are rust-colored, and the remiges and rectrices are black, some with white borders. The slim bill, the eyes, and the legs and feet are also blackish. Males and females cannot be distinguished by their looks; different behaviours in the breeding season is usually the only clue to the observer. Juveniles are even plainer in coloration, with buffy undertail coverts.
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They adopt various highly stereotyped and ritualised postures and associated plumage displays, which reveal prominent ocelli on remiges and rectrices. These behaviors are likewise used in self-defense. When utilised in pair-bonding behavior copulation may occur subsequent to lateral displays. Anterior displays are also performed which may include curious clicking and vibrating pulsations of feather quills created via stridulation.
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The plumage is predominantly white except on remiges, with a faint pink tinge on the neck and a yellowish base on the foreneck. The primary feathers are black, with white shafts at the bases, occasionally with paler tips and narrow fringes. The secondary feathers are also black, but with a whitish fringe. The upperwing coverts, underwing coverts, and tertials are white.
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In flight, the white underwing with black remiges of the adult are similar only to those of the American white pelican (P. erythrorhynchos), but the latter has white inner secondary feathers. It differs from the Dalmatian pelican in its pure white rather than greyish-white plumage, a bare pink facial patch around the eye, and pinkish legs. The spot-billed pelican (P.
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The underwing has grayish-brown remiges with white shafts to the outer primary feathers. The axillaries and covert feathers are dark, with a broad, silver-gray central area. The tail is dark gray with a variable silvery cast. The lower mandible is blackish, with a greenish-black gular pouch at the bottom for draining water when it scoops out prey.
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Very little information is known about the cryptic forest falcons breeding behavior. It is believed to nest in the wet season (December-May), as it molts remiges from April-August. This is also supported by possible damage from a nesting cavity in a female specimen collected in early June. It is presumed to breed in cavities as do other forest-falcons.
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The primary remiges are dark brownish-grey with off-white bars on the inner webs. One subspecies, M. r. zonothorax from the East Andean foothills, is polymorphic (at least in the northern part of its range), and also occurs in a brown morph, where most of the upperparts, head, and chest are brown or rufous instead of grey.Restall et al.
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An adult on twig The adults have a total length of approximately . They have grey upperparts and blackish remiges, but the colour of the remaining plumage depends on the subspecies. In the nominate subspecies and blythii, the underparts (incl. undertail) are rufous, but in nemoricola the underparts are whitish tinged rufous, especially on the flanks and crissum (the undertail coverts surrounding the cloaca).
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The upperwing of Charadrius ruficapillus shows dark brown remiges (flight feathers) and primary covert feathers with a white wingbar in flight. Its length is 14–16 cm and its wingspan is 27–34 cm; and weighs 35–40 g. Breeding plumage shows a red-brown crown and nape with black margins. Non-breeding plumage is duller and lacks the black margins.
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This is usually done high up in exposed locations. When they spot a predator they give alarm calls to alert the rest of the group to the type of threat. Pied babbler adults have a white head and body with dark brown rectrices and remiges. thumb Pied babbler fledglings form short-term associations with foraging adults, where they follow and beg to gain food.
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Altogether, the ability to fly and to perch was quite sophisticated for its age, to the detriment of terrestrial locomotion: the humerus was 1.56 times the length of the femur.Zhang et al. (2001), Lamanna et al. (2006) The holotype retains many feather impressions, though poorly preserved; remiges do not seem to have been preserved, and what feathers remain are apparently only body feathers, wing coverts and down.
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Females of this species have a slightly more olive nape and back than the males. The adult female's plumage is similar to the juvenile plumage; however, unlike adults, the wings are dull brown instead of black. In general, immature specimens have the hood; wingbars; remiges; and epaulets of adult specimens. The first-basic plumage retains the darker, greener coloration of the juvenile plumage, however.
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The wing coverts are a grizzled grey, and remiges mostly colourless grey. The male especially, has bright amaranth red underside plumage and bare, green gape and eye flanges. The female has brown face and chest plumage, blue skin orbiting the eyes and duller red plumage below. Immature birds resemble females, but have distinct white tips to the tertials (inner wing), and less distinct gape and eye flanges.
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Rhodospiza obsoleta The desert finch (Rhodospiza obsoleta), sometimes called Lichtenstein's desert finch, is a large brown true finch found in southern Eurasia. Its taxonomy is confused, and it has formerly been placed in Fringilla, Bucanetes, Carduelis and Rhodopechys. It has an average wingspan of . It has a stout black bill, black and white remiges and rectrices, and a slash of rosy-pink on each wing.
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The most critical period of feather molting runs from late August to early September. When viewed in flight, they have a misaligned or "moth-eaten" appearance and generally slower and more laborious travel. Their mobility is reduced due to the lack of several remiges or rectrices or these are not entirely renewed. Most of the red-winged blackbird have moved almost entirely by October.
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The major primary blankets are changed along with their respective primary springs. Unlike the major primary coats, the major secondary coats molt earlier than the secondary spruces. The molting of these feathers is rapid, with several of them at the same stage of development simultaneously. The progression of the molt in these feathers is from the outside to the inside, as in the secondary remiges.
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Legs are also yellow, though shade and brightness varies. Fledgling birds have undefined, fluffy light grey chests without scalloping. Immature birds can be identified in the hand by retained juvenile remiges and rectrices, which are more brownish. Yellow-throated miners are distinctive from the other miners by their clean white rump, instead of the continuous grey from the back that the noisy and black-eared miners have.
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The spotty bar over the greater wing coverts, narrow edging to the remiges and tips to the tertials are yellow. Additionally, the narrow yellow eyebrows extends as two parallel lines over the mantle. The belly is mainly pale yellow with black streaking to the flanks. Depending on subspecies, the throat ranges from red to orange, and the crown ranges from deep yellow over brownish-orange to reddish-orange.
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Museum specimen of juvenile The eagle is in length and has a wingspan of . Typical body mass is , with an occasional big female weighing up to . There is often a less obvious white patch on the upperwings, but a light crescent on the primary remiges is a good field mark. The white V mark on the rump is less clear-cut in adults than in the lesser spotted eagle (C. pomarina).
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Its face is blackish-brown and slightly darker than its head crown, which is a dull brown. The back is a paler chestnut-brown,pulling against the red rump and tail. The wing coverts are dark brown, with red or chestnut brown on the feather edges and tail end. The primaries are darker, the remiges black-brown with light edges and inner primaries marked with a short, light red wing bar.
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This indicates that the starling had strong neck and jaw muscles. According to Günther and Newton, the ulna of the Rodrigues starling was somewhat shorter than that of the hoopoe starling, measuring ; the humerus measured , and the keel on its sternum was a bit lower. It had strong quill knobs on the ulna, indicating that the secondary remiges were well developed. One coracoid measured in length, and one carpometacarpus was long.
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It has a total length of approximately 36–43 cm (14–17 in). It has a moderately sized black bill, a long tail and a mainly green plumage. The upperside of the remiges and primary coverts are blue, as indicated by its common name. The underside of the wings is yellowish, the tail-tip, crown and cheeks are bluish, and the tail-base and small belly-patch are red.
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The rockfowl's chin, throat, sides of the neck, and upper breast are all a pale grey. This bird is buffy lemon in colour on its lower breast, belly, flanks, thighs, and undertail coverts, though the flanks can sometimes appear to be greyish. The wing is grey, though the wing's remiges are black, forming a line between the lemon underparts and grey upperparts. Its legs and feet are silver-grey and muscular.
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Compared to the Eurasian magpie, it is somewhat stockier, with a proportionally shorter tail and longer wings. The back, tail, and particularly the remiges show strong purplish-blue iridescence with few if any green hues. They are the largest magpies. They have a rump plumage that is mostly black, with but a few and often hidden traces of the white band which connects the white shoulder patches in their relatives.
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Kn ("naked") is the most dominant of the allelic series which controls feather growth rate. It is a gene whose action on feather growth is quite extreme. Birds carrying this gene are nearly naked during juvenile life and in some cases females remain nearly naked well into adult life. One-day- old chicks lack primary and secondary remiges (they may appear as extremely small pinfeathers, but always shorter than the coverlets).
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In some passerines, filoplumes arise exposed beyond the pennaceous feathers on the neck. The remiges, or flight feathers of the wing, and rectrices, or flight feathers of the tail, are the most important feathers for flight. A typical vaned feather features a main shaft, called the rachis. Fused to the rachis are a series of branches, or barbs; the barbs themselves are also branched and form the barbules.
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The dusky parrot is approximately long. It has a medium-wide gray eye-ring (often fades to white in captivity), a splay of cream-white feathers on its upper neck, a pinkish-red tinge to the belly, and red undertail coverts. The remiges and tail are blue, with the latter red at the base. Overall, the dusky parrot is a dark brownish-gray (tending towards black in poor light) bird.
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It has a total length of about 24 centimeters (9½ in). The head and remiges are mainly rufous-chestnut, the underparts and back are buff, the wing-coverts are barred in black and buff and the chest and tail are uniform black. The male has a red malar and mottling on its crest. For comparison, the rufous- headed woodpecker is larger and has extensive black barring on the back and underparts.
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The common whitethroat (Curruca communis) is a common and widespread typical warbler which breeds throughout Europe and across much of temperate western Asia. This small passerine bird is strongly migratory, and winters in tropical Africa, Arabia, and Pakistan. This is one of several Curruca species that has distinct male and female plumages. Both sexes are mainly brown above and buff below, with chestnut fringes to the secondary remiges.
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A black breast band separates the lapwing's grey head and neck from the white underside. The wing coverts are brown. It has a variable but prominent white forehead patch similar to its near relative, the Senegal lapwing, but in contrast shows a prominent white wingbar in flight, bordered by black remiges. The two species are also separated by their respective habitat preferences, the Senegal lapwing preferring lower, mostly drier locations.
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The upper parts are mostly a darker fawn-brown, with the central rectrices and the primary remiges a little darker still; the head is colored like the underside, with a darker cap and light nape patch, somewhat reminiscent of some tits and chickadees, especially those from the genera Parus sensu stricto and Periparus. The bill, legs and feet are black. Males and females look alike.Londei (1998), del Hoyo et al.
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Bruce (1999) pp 34–75, Svensson et al. (1999) pp. 212–213 The bird's head and upper body typically vary between pale brown and some shade of grey (especially on the forehead and back) in most subspecies. Some are purer, richer brown instead, and all have fine black-and-white speckles except on the remiges and rectrices (main wing and tail feathers), which are light brown with darker bands.
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Males are highly vocal, and their loud, piercing whistle is frequently heard. It is strongly sexually dimorphic. Except for a bright yellow wing-speculum, males are superficially similar to the male common blackbird, while the far less conspicuous females are overall olive. The female resemble both sexes of the only other member of the genus, the grey- winged cotinga, but is larger, has a thicker bill, and yellowish-olive (not grey) remiges.
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The orange fruit dove (Ptilinopus victor), also known as flame dove, is a small, approximately long, short-tailed fruit-dove in the family Columbidae. One of the most colorful doves, the male has a golden olive head and elongated bright orange "hair-like" body feathers. The golden-olive remiges are typically covered by the long orange wing coverts when perched. The legs, bill and orbital skin are bluish-green and the iris is whitish.
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Male Common Shelduck in flight in England The common shelduck resembles a small short-necked goose in size and shape. It is a striking bird, with a reddish-pink bill, pink feet, a white body with chestnut patches and a black belly, and a dark green head and neck. The wing coverts are white, the primary remiges black, and the secondaries green (only showing in flight) and chestnut. The underwings are almost entirely white.
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The plumage is glossy black with a purplish sheen overall, though the rectrices and primary and secondary coverts have a greenish sheen and the remiges are a duller blackish-brown color showing reduced sheen. The female is less glossy than the male, and juveniles are brownish-gray with mottling below. The legs are black. The bill, which is black and shorter than the head, has a generally straight culmen, decurved toward the tip.
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Head The bird has a generally subdued coloration, with fine linear patterns of black, grey and brown. Its remiges however have vividly colored middle webs, which with wings fully spread show bright eyespots in red, yellow, and black. These are shown to other sunbitterns in courtship and threat displays, or used to startle potential predators. Male and female adult sunbitterns can be differentiated by small differences in the feather patterns of the throat and head.
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Measurements given here are for sharp-shinned hawk, but they are comparable for the remaining species.Raptors of the World by Ferguson-Lees, Christie, Franklin, Mead, and Burton. Houghton Mifflin (2001), Adults have short broad wings and a medium-length tail banded in blackish and gray with the tip varying among individuals from slightly notched through square to slightly rounded (often narrowly tipped white). The remiges (typically only visible in flight) are whitish barred blackish.
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Measurements given here are for the sharp-shinned hawk, but they are comparable for the remaining species.Raptors of the World by Ferguson-Lees, Christie, Franklin, Mead, and Burton. Houghton Mifflin (2001), Adults have short broad wings and a medium-length tail banded in blackish and gray with the tip varying among individuals from slightly notched through square to slightly rounded (often narrowly tipped white). The remiges (typically only visible in flight) are whitish barred blackish.
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The underside of the wing is pale rufous-buff, sometimes with some dark spotting on the underwing coverts. The tips of the primary remiges are barred with pale grey below, their bases are quite rufous. The iris is dark brown, the bill is black with a pale yellow cere; the feet are also pale yellow. Immature birds differ little from adults; they have lighter margins to the back feathers, producing a scalloped effect.
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Wing feathers Complete replacement of wing feathers takes about eight weeks. However, birds in their first year of age frequently retain some of the under-wing coverts and juvenile tertiary remiges after post-juvenile molting. Of seventy immature males examined during the last week of October, it retained 70% some older lower primary blankets. In most cases where partial replacement of the cover feathers occurs, it is the proximal covers that the bird retains.
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It has a total length of approximately . The neck is buffish, the upperparts are grey, the belly and flight feathers are black, and there is a large white patch in the wings. In flight, where the relatively short legs do not extend beyond the tail (unlike e.g. Eudocimus and Plegadis), the white patch forms a broad white band on the upperwing that separates the black remiges and the grey lesser wing-coverts.
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The upperparts, rectrices and undertail coverts are blackish- brown, as are at least the distal undersides of the remiges, but sometimes the entire feathers. The rest of the underparts are white, as is the head below eye level. The iris is dark, the feet are dull pink with a black wash and black toenails, and the bill is grey, darker towards the tip, and with a pinkish hue. Males and females look alike.
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The quail-plover is a small, short-tailed cursorial bird which looks a little like a miniature courser when on the ground. The upperparts are a sandy-rufous colour and the underparts mainly whitish. They show a distinctive wing pattern in flight when the contrast between the white primary coverts and the black with white-tipped remiges to form a distinct diagonal band on the upperwing. Its fluttering flight style is rather lark-like.
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Helmet vanga on nest, showing the massive bill Global distribution The helmet vanga is a large vanga, the second-largest species of vanga after the sickle-billed vanga. In length it measures , and it weighs . The most distinctive feature is the massive hooked bill, which is long and deep. The plumage of the head, neck, throat, breast and belly is a solid blue-black, as are the primary coverts and remiges of the wing.
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In worn individuals the bodily feathers of pale morph tawny eagles can appear almost whitish. Dark morph juvenile tawny eagles are generally light rufous to rufous brown with creamier lower back to upper tail coverts. Juveniles show thinly pale-tipped dark brown greater coverts and remiges while the tail is barred grey and brown usually with a narrow creamy tip. Dark morph juveniles may fade to pale buff or creamy often before molting into browner plumage.
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Buff-tailed coronet in Bellavista Cloud Forest Reserve, Ecuador The buff- tailed coronet (Boissonneaua flavescens) is a species of hummingbird from the family Trochilidae. It is found in Colombia, Ecuador, and Venezuela. Distinguishing characteristics are the short bill, glittering green coloration, and buffy color under the remiges (flight feathers) and rectrices (tail feathers). Like other hummingbirds, the buff-tailed coronet is often found in mid-level forest to canopy searching for flowers with nectar and some insects.
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Lilac-breasted rollers are not sexually dimorphic though males may be slightly larger than females. Juveniles, immatures and adults have the largest alula feather dark blue, but the primary coverts and rest of the alula azure. The proximal half of the remiges are also a brilliant azure, and the distal half black on the inner web, and dark purple blue on the outer webs. Juveniles have the throat and breast rufous-tawny with broad diffuse buffy-white streaks.
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The wings are barred with white, and the chin, throat and breast are in the male pure white, but of a bright reddish-orange in the female. The remiges are very short, rounded and much incurved, showing a bird of weak flight. The rectrices are very broad, the shafts stiff, and towards the tip divested of barbs. The population which is found locally in New Guinea is now generally considered a separate species, the Papuan logrunner, Orthonyx novaeguineae.
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Measurements given here are for the northern group, but they are comparable for the remaining species in the group.Raptors of the World by Ferguson-Lees, Christie, Franklin, Mead, and Burton. Houghton Mifflin (2001), Adults have short broad wings and a medium-length tail banded in blackish and gray with the tip varying among individuals from slightly notched through square to slightly rounded (often narrowly tipped white). The remiges (typically only visible in flight) are whitish barred blackish.
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The downy plumage of chicks is silky white on hatching, though it soon turns a smoky brown-grey. As in other sea eagles, remiges and rectrices of the first-year plumage are longer than those of adults. Juvenile plumage is largely a uniform dark soot- brown with occasional grey-brown streaking about the head and the neck, white feather bases, and light mottling on the rectrices. The tail of the immature eagle is white with black mottling distally.
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Nestlings are covered in pale buff down feathers, shading to whitish on the belly. Upperside pattern of male (presumably F. c./a. pallidus) wintering in Little Rann of Kutch, Gujarat, India The remiges are blackish, and the tail usually has some 3–4 wide blackish bands, too. Very light males only have faint and narrow medium-grey bands, while in the darkest birds the bands are very wide, so that the tail appears to have narrow lighter bands instead.
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Preserved feather traces in a fossil Zhenyuanlong suni. There is a large body of evidence showing that dromaeosaurids were covered in feathers. Some dromaeosaurid fossils preserve long, pennaceous feathers on the hands and arms (remiges) and tail (rectrices), as well as shorter, down-like feathers covering the body. Other fossils, which do not preserve actual impressions of feathers, still preserve the associated bumps on the forearm bones where long wing feathers would have attached in life.
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A rusty-red shoulder patch is just as characteristic when the bird is sitting with wings closed. The wings are dark above, admixed with grey near the bases of the blackish primary remiges. The underwing is whitish, with indistinct brownish barring on the underwing coverts that extends onto the flanks and thighs. The iris is hazel, the cere is pale green, the beak is black with a horn-colored base, and the feet are yellow with black talons.
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It has a narrow wing stripe, extending across the median and larger wing coverts, and often a bit onto the secondary remiges. Outer tail feathers never have white tips. Recent cladistic analysis of nDNA BRM15 intron-15 and mtDNA NADH dehydrogenase subunit 2 and ATP synthase F0 subunit 6 sequence data indicates that the Ethiopian boubou is a polyphyletic cryptic species complex, which was once lumped with the tropical boubou, black boubou, and East Coast boubou.
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In that same time rowers (remiges, pl. of remex) were trained on mock benches. Rowing was a skill requiring close coordination between the master, the steersman, the coxwain (Greek keleustes), stationed amidships, and the rowers, who must learn a repertoire of signals given by the coxwain. At the end of 60 days Scipio found himself admiral of a fleet of 160 new ships manned by newly trained oarsmen.The 160 is Florus', Book 2. Polybius has 120.
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Feathers of the back, breast, median wing coverts, rump and upper tail coverts are all iridescent green with a black base and some that are edged with bronze. The feathers on the head give a golden brown to golden green appearance, while the belly stands out as a brilliant red. The remaining feathers on the wings (remiges: primaries, secondaries; greater wing coverts), rectrices, and thighs are all black. The bill recalls the red of the belly, and yellows to the tip.
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The throat is unmarked white, the foreneck is white broadly streaked with pale brown, and the rest of the neck is buff with thin black barring. The breast and belly are white with broad pale brown streaks, while the back is buff, heavily streaked and barred with black. Rectrices are black in males and brown in females; the slate-grey remiges create a conspicuous two-toned effect in flight. The bill is stout and strong, yellowish overall with a dusky upper mandible.
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The capuchinbird is a large thick-set suboscine passerine with a relatively heavy bill. Adults weigh between and are typically around long, making it the largest suboscine passerine, apart from the Amazonian and long-wattled umbrellabirds – indeed, females average larger than any female umbrellabird. Its plumage is overall rich brown, approaching orange on the belly and undertail coverts, and the remiges and short tail are black. The most distinctive feature is its bare, almost vulture-like head covered in dull blue skin.
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Total length c. . As other members of the Pyrrhura picta complex, it is a long-tailed mainly green parakeet with a dark red belly, rump and tail-tip (tail all dark red from below), a grey- scaled chest, a whitish or dull yellow patch on the auriculars and bluish remiges. The forehead, ocular region and carpal edge are red in the nominate race (P. r. roseifrons). Its bare eye-ring is typically dark greyish (sometimes indistinct) bordered by yellowish-white.
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It is a slender, 15.5–17 cm long bird, with the long, constantly wagging tail characteristic of the genus Motacilla. The adult male in breeding plumage is basically grey or black above, with white on the remiges, and bright yellow below and on the entire head except for the black nape. In winter plumage, its yellow underparts may be diluted by white, and the head is brownish with a yellowish supercilium. Females look generally like washed-out versions of males in winter plumage.
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Courtship starts with the male chasing the female, the two hopping together through branches, and the male bobbing its head, bowing and giving a croaking call or low whistle. It makes gliding or descending display flights with the white rump spots exposed and producing mechanical noises with its remiges. The courtship climaxes with the male - wings drooped, tail fanned and rump feathers puffed up - giving repeated metallic or whistling calls. The couples may duet, but courtship feeding has not been observed.
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The crown is boarded by a black stripe extending from in front of the eyes until the gray auriculars. The tail and wings are blackish with the primaries margined slightly with a grayish external. There is a slight fulvous or tawny tint to the remiges most external parts. One important note is that the Cuzco brushfinch shows considerable variation with the intensity of gray in the underparts - some almost uniformly dark gray below and others that are pale gray with grayish-white abdomens.
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The female is mainly grey above, with a greyer head, and whitish with only light spotting. The Cyprus warbler's song is fast and rattling, and is similar to that of the Sardinian warbler. Together with Rüppell's warbler it forms a superspecies with dark throats, white malar streaks and light remiges fringes. This in turn is related to the species of Mediterranean and Middle East Curruca warblers that have a naked eye-ring, namely the subalpine warbler, Sardinian warbler and Menetries' warbler.
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At Biskeri Thatch (1,1000 ft ASL) in Kullu district, Himachal Pradesh (India) Hume's leaf warbler is one of the smallest "Old World warblers". Like most other leaf warblers, it has greenish upperparts and off- white underparts. With its long supercilium, crown stripe and yellow-margined tertial remiges, it is very similar to the yellow-browed warbler (P. inornatus). However, it has only one prominent light wing bar, just a faint vestige of the second shorter wing bar, and overall duller colours.
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Topaza pyra (mounted specimen) Topaza pyra can reach a body length of about . There is strong sexual dimorphism between the males and females. Both have a dark brown iris, but males are larger on average than females. Males of these brilliantly marked hummingbirds have a back, lower breast, upperwing-coverts, and outer webs of the innermost two remiges that are shining orange-red, becoming more orange on the belly, shading over the rump into the yellow-green/green uppertail-coverts.
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Shelducks are a group of large, often semi-terrestrial waterfowl, which can be seen as intermediate between geese (Anserinae) and ducks. They are mid-sized (some 50–60 cm) Old World waterfowl. The sexes are colored slightly differently in most species, and all have a characteristic upperwing coloration in flight: the tertiary remiges form a green speculum, the secondaries and primaries are black, and the coverts (forewing) are white. Their diet consists of small shore animals (winkles, crabs etc.) as well as grasses and other plants.
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The bill of the greater melampitta is also larger than that of the lesser melampitta, which is hooked. There is some variation in the family in the tails. The greater melampitta has specially strengthened remiges and retriges, which are often worn, a possible adaptation to its habit of roosting in limestone sinkholes. Those sinkholes are too deep and narrow to fly directly out and the tail may be used to help cling to the side of the hole as it exits, in the fashion of a woodpecker.
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Their proposed definition for the group was "the clade stemming from the first panavian with ... remiges and rectrices, that is, enlarged, stiff-shafted, closed-vaned (= barbules bearing hooked distal pennulae), pennaceous feathers arising from the distal forelimbs and tail".Gauthier, J. and de Queiroz, K. (2001). "Feathered dinosaurs, flying dinosaurs, crown dinosaurs, and the name 'Aves'". Pp. 7-41 in Gauthier, J. and L.F. Gall (eds.), New Perspectives on the Origin and Early Evolution of Birds: Proceedings of the International Symposium in Honor of John H. Ostrom.
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The name variable hawk is fully deserved, as both sexes occur in several morphs. Adults of all have a white tail with a contrasting black subterminal band and grey wings barred dark (in flight from below, the remiges appear whitish with fine barring and a broad black tip). The remaining plumage varies from very dark grey to whitish, and some individuals have reddish-brown to the underparts. Females usually have a reddish-brown back, which males usually lack, although at least some males also have this.
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It has conspicuous crimson bill ornaments--a round red knob with bony core adorns the maxilla base, while the cere extends apically at least halfway under this knob and below the mandible base forms a small fleshy wattle.del Hoyo (1994a) Females have black plumage just like the male, but their crissal area is reddish buff. In some, the remiges and sometimes the wing coverts have faint brownish marbling. Their bills and irides are also blackish, but their feet and legs are a greyish flesh color.
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Calls with the "spark" sounds at end The non-migratory genus Prinia shows biannual moult, which is rare among passerines. A moult occurs in spring (April to May) and another moult occurs in autumn (October to November). Biannual moult is theorized to be favoured when ectoparasite loads are very high, however no investigations have been made. Prinia socialis moults some remiges twice a year and is termed to have a partially biannual moult, however some authors describe P. socialis socialis as having two complete moults.
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Total length c. . As other members of the Pyrrhura picta complex, it is a long-tailed mainly green parakeet with a dark red belly, rump and tail-tip (tail all dark red from below), pale grey scaling to the chest, a whitish or dull buff patch on the auriculars and bluish remiges. The cheeks and crown are dark dusky-maroon (often appears almost blackish). Unlike other members of the P. picta complex, it lacks any bright red or blue to the head (but see Taxonomy).
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The structures were first noted by P. L. Sclater in 1860, and the sound production adaptations were discussed by Charles Darwin in 1871 Like several other manakins, the club-winged manakin produces a mechanical sound with its extremely modified secondary remiges, an effect known as sonation. The manakins adapted their wings in this odd way as a result of sexual selection. In manakins, the males have evolved adaptations to suit the females' attraction towards sound. Wing sounds in various manakin lineages have evolved independently.
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The remiges and primary coverts are blue and the long, pointed tail has a red base, a narrow green center and a blue tip. The underside of the tail and flight feathers are greenish-yellow, similar to that of several other small macaws such as the blue-winged and red-bellied macaw. The legs are a dull pinkish color, and the iris is reddish to dull yellow. It has extensive bare white facial skin and the heavy bill is black, often tipped pale grey.
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Flight feathers also have the ability to twist their rachis inside the pulp cavity, allowing the barbules of the feather to interlock on the down stroke, decreasing the amount of air flow between the feathers, creating a more efficient lift. The flight feathers of the wing are remiges; those located on the tail are rectrices. Flight feathers are separated into three separate groups—primary, secondary and tertiary. The primaries are at the far end of the wing and provide the forward thrust during takeoff and flight.
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The males also have an orange-tipped black tail, black, orange and white wings, a bright orange bill, an orange iris yellowing as it nears its outer edge, and silky- orange filamentous feathers of the inner remiges. Both sexes also have orange legs and skin. The less conspicuous female is dark brownish-grey overall and has a yellow-tipped black bill, a duller orange iris, and a smaller crest. One-year-old juvenile males look similar to an adult female, but has orange speckles over their bodies.
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The edges of black flight feathers and greater primary and secondary coverts are bright yellow, as are those of the tail feathers, which also have white tips. The striking colour contrast in the feathers of the remiges and rectrices is one of the species' most distinctive traits, along with its pink bill, which has a grey tip. Feet and legs are also grey, while the irises of the eyes are reddish-brown . Females are slightly smaller than the males and are paler in colour, with fewer spots running down the flanks.
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Immature birds resemble the adult females, but are duller and have hardly any white at the bill base. Both sexes have white secondary remiges, a blue-grey bill with a black "nail" at the tip and grey feet; the drakes have a bright yellow iris, while that of females is orange or amber and that of immatures is brown. Downy hatchlings look much like those of related species, with dark brown upperparts and pale buff underparts, chin, supercilium and back spots. These birds are not very vocal, at least compared to dabbling ducks.
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The throat, cheeks and belly often have a bluish tinge. As most members of the genus Amazona, it has broad dark blue tips to the remiges and a red wing- speculum. Its beak is horn coloured. In its range the yellow shoulder patch and extensive yellow on the head distinguish the yellow-shouldered amazon from other Amazona species, which have red or orange on the shoulder and less yellow on the head (the orange-winged amazon, which has as much yellow to the head as some yellow-shouldered amazons, has a blue ocular region).
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It is considerably darker than other adult Aquila eagles in central Eurasia. Furthermore, all other Eurasian eagles in their range lack the white spots on the wing mantle and greyish under-tail. Given reasonable views, the juvenile imperial eagle is no less distinctive, with its unique tawny-buff covered in brownish streaking, a colour combination not seen in other species. From a distance, the juvenile may give the impression of a dark mantle and chest band with very pale rear body and a blackish tail and remiges against strikingly pale primary wedges.
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Young birds are white for about two- thirds of their tail length, ending with a broad, black band. Occasionally, juvenile eagles have white patches on the remiges at the bases of the inner primaries and the outer secondaries, forming a crescent marking on the wings which tends to be divided by darker feathers. Rarely, juvenile birds may have only traces of white on the tail. Compared to the relatively consistently white tail, the white patches on the wing are extremely variable; some juveniles have almost no white visible.
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The adults' upper parts and tail are glossy blue-black except for concealed white spots on the rump, visible only when the wings are spread and the rump feathers are erected. The underparts are white, in some populations with a buffy or pinkish tinge on the breast and flanks, which is not always noticeable except in good light. The wings of most subspecies have a white stripe on the wing coverts, in some extending onto the secondary remiges. The tips of the outer tail feathers can be white in some subspecies.
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Dorsal view of Mangrove robin showing white in outer rectrices The mangrove robin has an average weight of for males and for females. Their wingspan differs between subspecies – the leucura subspecies have spans of to for males and to for females, while the alligator subspecies have spans of to for males and to for females. For cinereiceps, male birds have wingspans of to long; on the other hand, female wingspans are to long. They feature a "dull pale bar" at the bottom of their remiges, although this is not very noticeable.
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The progression of the molting of the lower middle secondary blankets is from the outside to the inside, while that of the lower middle primary mats seems to be irregular or almost simultaneous. The medial lower coverts molt before the primary remiges VIII and IX. The major lower primary coverts and major lower secondary coverts molt last. The progression of the molting of these last feathers is the same as in the primary and secondary sprouts, that is, from the inside out and from the outside in, respectively.
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Upperwings are also dark grey-brown, but with prominent white shafts and narrow rufous-brown fringes to the secondary coverts and tertials; and fine light brown edges to the other remiges, producing rufous-brown patches when the wing is folded. The underbody is white with buff-brown wash on the flanks through to the legs and underside of tail. The bill is light grey to blue-grey with a darker grey culmen, and the iris is dark to olive-brown. The legs and feet are purplish to dark grey.
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Together with the Cyprus warbler it forms a superspecies with dark throats, white malar streaks and light remiges fringes. This in turn is related to the species of Mediterranean and Middle East Sylvia warblers that have a naked eye-ring, namely the subalpine warbler, Sardinian warbler and Ménétries's warbler. Both groups have a white malar area, but this may not form a clear streak in the latter group; above the white, the heads of males are uniformly dark.Helbig, A. J. (2001): Phylogeny and biogeography of the genus Sylvia.
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They start to grow feathers 5 days or so after hatching, starting with the remiges; the rectrices begin to emerge about 3 days later. The young are fed 1-2 times per hour on average, and the female spends about half of the day brooding and feeding her offspring, and the other half flying around and feeding. The young fledge after 22–24 days but still return to the nest to sleep and be brooded for some more days; they are independent some 2–3 weeks after fledging.
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Secondly, serrated edges on the leading edge of owls' remiges muffle an owl's wing beats, allowing an owl's flight to be practically silent. Some fish-eating owls, for which silence has no evolutionary advantage, lack this adaptation. An owl's sharp beak and powerful talons allow it to kill its prey before swallowing it whole (if it is not too big). Scientists studying the diets of owls are helped by their habit of regurgitating the indigestible parts of their prey (such as bones, scales, and fur) in the form of pellets.
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2, A field guide. Christopher Helm. Juvenile hook-billed kite (Chondrohierax uncinatus) are also potentially confusable with juvenile ornates but the kite is much smaller and more dumpily built with more paddle- shaped wings, a squarer tail, with clearer bars on remiges and rectrices and bare tarsi. Another kite, the gray-headed kite (Leptodon cayanensis) can be considered similar in plumage in its adult plumage to the juvenile ornate but it is rather smaller with very different shape in all respects (especially in its small, pigeon-like head), completely different underwing pattern and unmarked body but for grey crown and nape.
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In common with other honeyeaters, the red wattlebird has a long, specialized tongue to extract nectar from flowers. The tongue can extend well beyond the tip of the bill, and is divided at the end to form a brush-like structure with over a hundred bristles that soak up nectar by capillary action. The red wattlebird begins moulting after the breeding season, starting with the primary flight feathers in November or December, and finishing between the following March and May. The feathers of the breast, back, median and lesser covert feathers are moulted before those of the crown, remiges, and rectrices.
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Consequently, it is considered to be of least concern by BirdLife International and IUCN. This long-tailed species is generally green in color (both lutino and blue mutations are rare, but do exist in captivity) with a gray-brown head, a blue-tipped tail, and remiges that are dark gray from below, mainly blue from above. The bill is black, and it has a broad, bare, white (sometimes yellow-tinged) eye-ring. With a typical length of 25–28 cm (10–11 in) and a weight around 100 grams, it is slightly smaller than the sun conure.
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The white feathers of the head and rump have concealed dark brown bases, while those of the mantle, back, tail rectrices and tail coverts have dark brown shaft bases. The two long tail feathers are orange or red with white bases for around a tenth of their length, and can be hard to see when the bird is flying. The white wings are marked by dark chevron-shaped patches on the tertials, and the dark shafts of the primary flight feathers are visible. The pink tinge is often more pronounced in the remiges of the upper wing.
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Patagium impressions of Confuciusornis (a, b) compared to the patagium of a chicken (c, d) The wing feathers of Confuciusornis were long and modern in appearance. The primary wing feathers of a 0.5-kilogram individual reached 20.7 centimetres in length. The five longest primary feathers (remiges primarii) were more than 3½ times the length of the hand and relatively longer than those of any living bird, while the secondary feathers of the lower arm were rather short by comparison. The outermost primary was much shorter than the second outermost primary, creating a relatively round, broad wing.
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60–61, 150–151 Males and females are about the same size, and do not differ conspicuously in appearance except by direct comparison. In the female the underparts are greyer and are usually visibly barred greyish-brown, and the white wing and tail markings are characteristically less in extent (though this is rarely clearly visible except in flight). Fledged young birds are heavily tinged greyish-brown all over, with barring on the upperside and indistinct buffy-white markings. The tips of the tertiary remiges and the wing coverts are also buffy, with a black band in the latter.
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Egg of a Lagopus The three species are all sedentary specialists of cold regions. Willow ptarmigan is a circumpolar boreal forest species, white-tailed ptarmigan is a North American alpine bird, and rock ptarmigan breeds in both Arctic and mountain habitats across Eurasia and North America. With the exception of the red grouse, all have a white winter plumage that helps them blend into the snowy background. Even their remiges are white, while these feathers are black in almost all birds (even birds that are predominantly white, such as the Bali myna) because melanin makes them more resilient and thus improves flight performance.
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The plumage is almost entirely bright white, except the black primary and secondary remiges, which are hardly visible except in flight. From early spring until after breeding has finished in mid-late summer, the breast feathers have a yellowish hue. After moulting into the eclipse plumage, the upper head often has a grey hue, as blackish feathers grow between the small wispy white crest. The bill is huge and flat on the top, with a large throat sac below, and, in the breeding season, is vivid orange in color as are the iris, the bare skin around the eye, and the feet.
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Adult eagles that do show a dark-barred greyish primary patch usually have that confined to a wedge-shape on inner primaries though can sometimes be rather more prominent. Below adults show dark-barred grey flight feathers and tail with the broad blackish trail edges and wing ends being rather distinctive; the wing linings are often slightly paler to darker than remiges and often with an obscure remnant of broken paler central band. Juveniles are quite distinctive in flight if seen in reasonable view. Above, juveniles are pale greyish-brown to yellow-brown about the body and forewing-coverts, have a broad whitish U above the tail.
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Illustration by John Gould (probably from stuffed specimens) Illustration in habitat by Gustav Mützel The tooth-billed pigeon is a medium-sized, approximately 31 cm long, dark pigeon with reddish feet and red bare skin around the eye. The underparts, head and neck are greyish with a slight blue-green iridescence, and the tail, wings- coverts and tertials are rufous chestnut, while the remaining remiges are blackish. It has a large, curved, and hooked bright red bill with tooth-like projections on the lower mandible. Both sexes are similar, but the juvenile is duller with a browner head, with a black bill with only the base a pale orange.
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The dull-mantled antbird is long and weighs around . Overall, these birds look essentially blackish grey in the front half and dark reddish brown in the hind part, with a black wing-patch with white spots right where the two main colors meet. But in the dusky forest understory, the birds may appear all-black, with only the white spotting standing out.Zimmer & Isler (2003a) The plumage of the male is blackish grey on the head, neck, upper mantle and on the underside up to the upper belly, and reddish brown on most of the remaining upperparts and underparts; remiges and rectrices are somewhat darker, with dark reddish brown edges.
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Though mimids were widely considered Turdidae until the 1850s, this was not any more correct than treating them as Old World flycatchers, as these three families are distinct lineages of the superfamily Muscicapoidea. In the mid-20th century, the Turdidae and even most of the Sylvioidea were lumped in the Muscicapidae—but the Mimidae were not. Lastly, the smaller gray catbirds from Bermuda, which have proportionally narrow and shorter rectrices and primary remiges, were described as subspecies bermudianus ("from Bermuda") by Outram Bangs in 1901. But this taxon was never widely accepted, and today the gray catbird is generally considered monotypic as a species, too.
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Juvenile males shed more remiges than females when moulting into adult plumage. East of the 106th meridian west, birds moult strongly before breeding and replace another quantity of feathers afterwards, and post-juvenal moult does not differ significantly between the sexes. However, this seems dependent on the differing rainfall regimes; simply put, birds at least anywhere in the North American range moult most of their plumage at the end of the dry season and may replace more feathers at the end of the wet season. Considered a Species of Least Concern by the IUCN due to its vast range, it nonetheless seems to be declining locally.
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A fairly pale adult female (note brown remiges and yellow eye) winters near Hodal (Faridabad district, Haryana, India) Adult male (front), juvenile (behind) and adult female (back), illustration from 1899 The western marsh harrier is in length, has a wingspan of and a weight of in males and in females. It is a large, bulky harrier (larger than other European harriers but slightly smaller than the eastern marsh harrier) with fairly broad wings, and has a strong and peculiar sexual dichromatism. The male's plumage is mostly a cryptic reddish-brown with lighter yellowish streaks, which are particularly prominent on the breast. The head and shoulders are mostly pale greyish- yellowish.
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But see Bolle (1857) who records the South African birds occurring as far north as Gabon. H. meadewaldoi had a glossy black coloration overall save for the whitish underwing bases of the primary remiges' inner webs, but this may have not been present in worn plumage which also was duller. Its bill, laterally compressed and with a blunt, lighter tip, and a narrow naked ring around the red eye were reddish orange, and the legs and feet were dark pink with ivory-colored nails. As usual in oystercatchers, it had no hallux and the second and third toes were connected by a small web.
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The remiges, rectrices, alula, medium, greater and primary coverts are a dark grey colour with a brown tinge, with bright greyish-green outer edges. The juvenile resembles the adults, but has looser, fluffier plumage and, possibly as a result of this, slightly less yellow underparts. It has marginally whiter and more clear-cut pale tips to the greater coverts, and probably slightly more green on the anterior part of the lateral crown-stripes. “In plumage, the Limestone Leaf Warbler appears to be largely indistinguishable from P. ricketti, although as a result of the small number of specimens available for the Limestone Leaf Warbler and their poor quality, detailed comparisons are difficult to make.
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The wings and the square-tipped tail are black, with blue iridescence on the primary and secondary wing-coverts, and a striped patch formed by the light grey outer web of the tertiary remiges, with the grey scapulars sometimes dropping over the wings to form a grey shoulder patch. The uppertail coverts are likewise black, quite long, and have white tips that may have a signal function as they are sometimes prominently presented by the birds. The belly is white, and the throat has a patch of cherry red that varies in extent, in some birds reaching onto the breast. The feet and toenails are dull pink, and the iris is dark yellow.
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It had teeth only in the front of the jaws, with unusually long front teeth angled forward, a feature only seen in Masiakasaurus among other theropods. The rest of the skeleton bore an overall similarity to the possibly closely related Scansoriopteryx, including a hip configuration unusual among other dinosaurs: the pubis was shorter than the ischium, and the ischium itself was expanded towards the tip. The tail of Epidexipteryx also bore unusual vertebrae towards the tip which resembled the feather-anchoring pygostyle of modern birds and some oviraptorosaurs. Restored skull Epidexipteryx appears to have lacked remiges (wing feathers), though based on the related Yi, it may have possessed some sort of membrane wing to allow gliding.
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The adult males (drakes) in alternate plumage have a black, iridescent head and a small tuft at the hindcrown, a black breast, a whitish-grey back and wings with darker vermiculations and black outer and greyish-brown inner primary remiges. The underparts are white with some olive vermiculations on the flanks, and the rectrices and tail coverts are black. Adult females (hens) have a white band at the base of the bill, often a lighter ear region, and are otherwise dark brown all over, shading to white on the mid-belly. Drakes in eclipse plumage look similar, but with a very dark head and breast, little or no white on the head and usually some greyish vermiculations on the wings.
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When quill knobs are present, this is considered a strong indication that the animal had long remiges on the wings. Since quill knobs are rare in the fossil record, paleontologists mainly relied on phylogenetic bracketing to determine if a species was likely to have had wing feathers - if a relative on a "higher" branch of the evolutionary tree had the feathers, and one on a "lower" branch down had them also, then a species in the middle position likely did as well. Dakotaraptor’s quill knobs show that the animal unequivocally had prominent wing feathers, making it the largest dromaeosaurid with confirmed plumage of that type. The quill knobs of Dakotaraptor have a diameter of about , which shows that these feathers were rather large.
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The only differences between the Miyako- jima bird and males of the Guam kingfisher (the nominate subspecies of the Micronesian kingfisher; presently only surviving in captivity) are the former's lack of a black nape band and the red feet (black in Guam birds). The bill color is unknown due to damage to the specimen, and supposed differences in the proportion of the remiges are almost certainly an artifact of specimen preparation. Indeed, the specimen was not recognized as distinct until some 30 years after its collection. If the bird was indeed a resident of the Miyako group (and as there was better habitat on neighboring Irabu-jima, it is probable that it would have been found there too), it became extinct in the late 19th century.
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Reconstructions usually depict it with feathers, as Chatterjee originally interpreted structures on the arm to be quill knobs, the attachment point for flight feathers found in some modern birds and non-avian dinosaurs. However, re-evaluation of the fossil material by subsequent authors such as Lawrence Witmer have been inconclusive regarding whether or not these structures are actual quill knobs. In his 1997 account, Chatterjee infers the presence of feathers from alleged quill knobs on the badly smashed ulna and metacarpals III and IV, and infers the presence of remiges from such structures (though he does caution that this is uncertain). As is the case with the alleged quill knobs on the ulna, the metacarpal structures appear to be attributable to post-mortem damage.
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It is notable that at least the primary remiges, and often the other flight feathers too, are typically black in birds – even if the entire remaining plumage is completely white, as in some pelicans or in the Bali starling (Leucopsar rothschildi). This is due to the fact that melanins will polymerize, making all-black feathers very robust; as the largest forces encountered by bird feathers affect the flight feathers, the large amount of melanin gives them better resistance against being damaged in flight. In soaring birds as dependent on strong winds as the bony-toothed birds were, black wingtips and perhaps tailsShort-tailed waterfowl and "higher waterbirds" often have light or white tails: del Hoyo et al. (1992) can be expected to have been present.
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Primary remiges are one of the first feathers to molt. The molt of these feathers proceeds regularly from the innermost primary—primary I—to the outermost—primary IX. By October 1, most birds have either acquired the three new external primaries—VII, VIII, and IX—or they are at some advanced stage of development. The average dates for completion of the development of the new primary springs are: August 15 for primary I; September 1, primary II-IV; September 15, primary V and VI; and October 1, primaries VII-IX. The molting of the secondary rémiges begins with the most external—secondary I— and proceeds inward to secondary VI. The secondary I sheath appears around the same time that all the secondary covers have been replaced and rarely before mid-August.
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It is a robust bird, with a stocky, strong beak and a long tail. It measures some 15 cm long. The scientific name emphasizes the absence of the flamboyant livery typical of many Australian species, as the brown tones predominate in the plumage of this bird; the upper part of the body (nape, back, wings and tail) is in fact a deep brown color, with a tendency to darken on the tail, while the ventral area (cheeks, throat, chest, abdomen and hips) is beige color that turns towards white in the central part of the belly and on the undertail. The bird distinguishes for its fine zebra like colouring, which is white and present on the eyebrows, neck, chest, hips and tail, while on the remiges are two rows of white spots.
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It differs from the contemporary Pseudocrypturus by a shorter skull - in Calciavis the skull is shorter than the humerus, while in inverse happens in Pseudocrypturus -, as well as a proportionally narrower coracoid shaft and longer tarsometatarsus, from Lithornis promiscuus in aspects of the ischium, and from Paracathartes in a less curved and more gracile scapular blade. Feather imprints show abundant plumage with long primaries and remiges. In AMNH 30578 most of it is damaged due to post-mortem decomposition, with disorganised patches in the pectoral and pelvic region and the left wing traces and impressions being damaged, but the right wing is mostly intact, even showing evidence of barbules; in AMNH AMNH 30560 a wing is similarly well preserved. It is unclear if it had a tail, as the left wing feathers block the caudal region in AMNH 30578, but other lithornithids lack tail feathers.
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He concluded that his interpretation was the ulnar bumps were part of the crest on the anterior side of the ulna perhaps for insertion of the musculature that connected the ulna and radius, pointing out that Balaur has a crest on the anterior side of its ulna which is homologous with that of Concavenator and has no relation with remiges. However, in 2018 Cuesta Fidalgo published her doctorate thesis on the anatomy of Concavenator which argued that the ulna was preserved in lateral view, meaning that the ulnar bumps were positioned posterolaterally rather than anterolaterally as Cau and Mortimer claimed. Cuesta Fidalgo noted that the proximal part of the ulna is affected by fracturing and abrasion, and certain features would have shifted compared to their position in the bone when the animal was alive. For example, in the fossil, the lateral process of the ulna is positioned further posteriorly than the ulnar bumps.
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Swainson's hawks are distinctly darker on the wing and ferruginous hawks are much paler winged than typical red-tailed hawks. Pale morph adult ferruginous hawk can show mildly tawny-pink (but never truly rufous) upper tail, and like red-tails tend to have dark markings on underwing-coverts and can have a dark belly band but compared to red-tailed hawks have a distinctly broader head, their remiges are much whiter looking with very small dark primary tips, they lack the red- tail’s diagnostic patagial marks and usually (but not always) also lack the dark subterminal tail-band, and ferruginous have a totally feathered tarsus. With its whitish head, the ferruginous hawk is most similar to Krider's red- tailed hawks, especially in immature plumage, but the larger hawk has broader head and narrower wing shape and the ferruginous immatures are paler underneath and on their legs. Several species share a belly band with the typical red-tailed hawk but they vary from subtle (as in the ferruginous hawk) to solid blackish, the latter in most light-morph rough-legged buzzards.
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Anton III Wierix 1606). The ten quaternions are shown underneath the emperor flanked by the prince-electors (Archbishop of Trier, Archbishop of Cologne, Archbishop of Mainz; King of Bohemia, Count Palatine, Duke of Saxony, Margrave of Brandenburg). A " Quaternion Eagle" (each quaternion being represented by four coats of arms on the imperial eagle's remiges) Hans Burgkmair, c. 1510. Twelve quaternions are shown, as follows (eight dukes being divided into two quaternions called "pillars" and "vicars", respectively c.f. Christian Knorr von Rosenroth, Anführung zur Teutschen Staats-Kunst (1672), p. 669.): Seill ("pillars"), Vicari ("vicars"), Marggrauen (margraves), Lantgrauen (landgraves), Burggrauen (burggraves), Grauen (counts), Semper freie (nobles), Ritter (knights), Stett (cities), Dörfer (villages), Bauern (peasants), Birg (castles). The so- called imperial quaternions (German: Quaternionen der Reichsverfassung "quaternions of the imperial constitution"; from Latin quaterniō "group of four soldiers") were a conventional representation of the Imperial States of the Holy Roman Empire which first became current in the 15th century and was extremely popular during the 16th century.Hans Legband, "Zu den Quaternionen der Reichsverfassung", Archiv für Kulturgeschichte 3 (1905), 495–498. Ernst Schubert, "Die Quaternionen", Zeitschrift für historische Forschung 20 (1993), 1–63.
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The silvery pigeon is not distinguishable from the pied imperial pigeon at a distance, although this is not necessarily true vice versa; as the pied imperial pigeon can vary between a pale grey, pure white and even yellowish colouration, it is often possible to tell that a bird is not a C. argentina. At close quarters, the silvery pigeon may be recognized by a few characteristics: The plumage is always a pale silvery grey, with black remiges and ends of the tail feathers; there may be a slight greenish sheen on the feathers of the backsides of the neck. The black part of the tail is equal in length in all feathers, whereas it forms a black triangle pointing headwards on the underside of the pied imperial pigeon's tail. Most distinguishing characteristics are located on the head, which is shaped differently, with a sloping forehead (rounded in the PIP), conspicuous dark red or purplish eye-wattles (none in the PIP) and eyes, and a bill that is darker at the base (lighter at the base in the PIP), being dusky purple with a pale apple-green tip.
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